Flowers and fruit on female plant. Crystal Cove State Park, Laguna Beach, CA 8/6/12. © Peter J. Bryant

Fourwing Saltbush

Atriplex canescens

Chenopodiaceae

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var. canescens, Nr. La Novia Ave., E Valley Rd., San Juan Cap., 3-1-15. © Ron Vanderhoff.
var. canescens, pistillate (female) L, staminate (male) R, Nr. La Novia Ave., E Valley Rd., San Juan Cap., 3-1-15. © Ron Vanderhoff.

Leaves. Crystal Cove State Park, Laguna Beach, CA 8/6/12. © Peter J. Bryant
Bluffs, SW end of Linda Ln, San Clmte, 10-29-11. © Ron Vanderhoff.
Female (pistillate) plants

Flowers on female plant. Crystal Cove State Park, Laguna Beach, CA 8/6/12. © Peter J. Bryant
Flowers on female plant. Crystal Cove State Park, Laguna Beach, CA 8/6/12. © Peter J. Bryant

Female plant. Crystal Cove State Park, Laguna Beach, CA 9-27-07. © Joan Avise
Flowers and fruit on female plant. Crystal Cove State Park, Laguna Beach, CA 8/6/12. © Peter J. Bryant

Fruit on female plant. Crystal Cove State Park, Laguna Beach, CA 8/6/12. © Peter J. Bryant
Mature fruit on female plant. Crystal Cove State Park, Laguna Beach, CA 9-27-07. © Joan Avise
Male (staminate) plants

Male plant. Big Canyon, Upper Newport Bay, Newport Beach, Orange County, CA. 7-10-10. © Peter J. Bryant
Male plant. Big Canyon, Upper Newport Bay, Newport Beach, Orange County, CA. 7-10-10. © Peter J. Bryant

Immature inflorescences on male plant. Crystal Cove State Park, Laguna Beach, CA 8/6/12. © Peter J. Bryant
Male inflorescences. Crystal Cove State Park, Laguna Beach, CA 8/6/12. © Peter J. Bryant

Immature inflorescence on male plant. Crystal Cove State Park, Laguna Beach, CA 8/6/12. © Peter J. Bryant

Male inflorescence. Crystal Cove State Park, Laguna Beach, CA 8/6/12. © Peter J. Bryant

Male plant. Big Canyon, Upper Newport Bay, Newport Beach, Orange County, CA. 7-10-10. © Peter J. Bryant
Salt Creek mouth, beach to 1 mi NW, 7-19-12. © Ron Vanderhoff.

Salt Creek mouth, beach to 1 mi NW, 7-19-12. © Ron Vanderhoff.
Salt Creek mouth, beach to 1 mi NW, 7-19-12. © Ron Vanderhoff.
Associated Insects

Saltbush Gall Midge,
Asphondylia (solidaginis?)
Parasitoid Torymus atriplicis, female
Parasitoid PBTor2, Female

Stem Gall induced by Saltbush Gall Midge. Big Canyon, Newport Beach, CA. 7/10/10. © Peter J. Bryant.